As America searched for their ideal tank destroyer vehicle, evaluation of a "Cook Interceptor" 8x8 vehicle proved to be successful enough to be considered for a tank destroyer variant holding a 3-inch gun. Initially produced in November 1942 as the 3-inch Gun Motor Carriage T55, the test results and requirements were incorporated into the design of the second unit. The second unit was produced in April 1943 as the 3-inch Gun Motor Carriage T55E1. While the wheeled performance was considered satisfactory, the vehicle's mobility was unsatisfactory against the requirement that the vehicle be as mobile and manoeuvrable as a tank. The vehicle was not accepted into service and was sent to Aberdeen Proving Grounds in August 1943.

Introduced during Update "Hot Tracks" as a reward for Battle Pass: Season II, "Steel Centurion", the T55E1 is a very mobile tank destroyer. The wheeled design brings limitations in its cross-country mobility and turning ability, but gives the T55E1 unprecedented forward and reverse speed for a 3-inch gun platform. The 3-inch gun, being so far forward, gives the luxury of presenting as small a target as possible when exposed towards the enemy to fire, especially when utilizing the vehicle's good gun depression. The T55E1 is a very different tank destroyer compared to the usual American line-up, but its gameplay focusing on its low profile and wheeled car mobility gives the T55E1 great hit-and-run characteristics to quickly get in and out of dangerous situations.